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636-0009-00L 6 Credits MSC D-BSSE , D-INFK , D-BIOL , D-MATH
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Evolutionary Dynamics

Lecturers & Examiners: Prof. Dr. Niko Beerenwinkel
VVZ CR n/a

Last Updated: 2026-02-05 15:36:20

Abstract

Evolutionary dynamics is concerned with the mathematical principles according to which life has evolved. This course offers an introduction to mathematical modeling of evolution, including deterministic and stochastic models.

Objective

The goal of this course is to understand and to appreciate mathematical models and computational methods that provide insight into the evolutionary process.

Content

Evolution is the one theory that encompasses all of biology. It provides a single, unifying concept to understand the living systems that we observe today. We will introduce several types of mathematical models of evolution to describe gene frequency changes over time in the context of different biological systems, focusing on asexual populations. Viruses and cancer cells provide the most prominent examples of such systems and they are at the same time of great biomedical interest. The course will cover some classical mathematical population genetics and population dynamics, and also introduce several new approaches. This is reflected in a diverse set of mathematical concepts which make their appearance throughout the course, all of which are introduced from scratch. Topics covered include the quasispecies equation, evolution of HIV, evolutionary game theory, birth-death processes, evolutionary stability, evolutionary graph theory, somatic evolution of cancer, stochastic tunneling, cell differentiation, hematopoietic tumor stem cells, genetic progression of cancer and the speed of adaptation, diffusion theory, fitness landscapes, neutral networks, branching processes, evolutionary escape, and epistasis.

Resources

Lecture Notes

No.

Literature

- Evolutionary Dynamics. Martin A. Nowak. The Belknap Press of Harvard University Press, 2006. - Evolutionary Theory: Mathematical and Conceptual Foundations. Sean H. Rice. Sinauer Associates, Inc., 2004.

General Information

Language
English
Levels
MSC
Frequency
Yearly recurring

Examination

Type
session examination
Mode
oral 20 minutes
Repetition possible only with re-enrollment, including projects.The final grade is 70% oral session examination and 30% project. The practical projects are compulsory continuous performance assessments (60 hours of work, 2 credits) of the course. The project has to be re-run in case of a repetition.

Course Components

Type Title Time & Place Hours
lecture Evolutionary Dynamics
Attention: lecture starts Thursday, September 24 Lecture: 9-11, Tutorial: 11-12h The lecturers will communicate the exact lesson times of ONLINE courses.
  • Thu 09:00-11:00 (ON LI NE)
2 h weekly
exercise Evolutionary Dynamics
The lecturers will communicate the exact lesson times of ONLINE courses.
  • Thu 11:00-12:00 (ON LI NE)
1 h weekly
independent project Evolutionary Dynamics
Project Work (compulsory continuous performance assessment), no fixed presence required.
No time listed 2 h weekly

Offered In